Accounts of unusual sexual activities among penguins, observed a century ago by a member of Captain Scott's polar team, are finally being made public.
Details, including "sexual coercion", recorded by George Murray Levick were considered so shocking that they were removed from official accounts.
However, scientists now understand the biological reasons behind the acts that Dr Levick considered "depraved".
The Natural History Museum has published his unedited papers.
Mr Levick, an avid biologist, was the medical officer on Captain Scott's ill-fated Terra Nova expedition to the South Pole in 1910. He was a pioneer in the study of penguins and was the first person to stay for an entire breeding season with a colony on Cape Adare.
He recorded many details of the lives of adelie penguins, but some of their activities were just too much for the Edwardian sensibilities of the good doctor.
He was shocked by what he described as the "depraved" sexual acts of "hooligan" males who were mating with dead females. So distressed was he that he recorded the "perverted" activities in Greek in his notebook.
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Only two of the original 100 copies of Mr Levick's account survive. Mr Russell and colleagues have now published a re-interpretation of Mr Levick's findings in the journal Polar Record.
Mr Russell described how he had discovered one of the copies by accident.
"I just happened to be going through the file on George Murray Levick when I shifted some papers and found underneath them this extraordinary paper which was headed 'the sexual habits of the adelie penguin, not for publication' in large black type.
"It's just full of accounts of sexual coercion, sexual and physical abuse of chicks, non-procreative sex, and finishes with an account of what he considers homosexual behaviour, and it was fascinating."